Jonathan was one of the members of Rat Patrol Team 01 involved in the Guns of the Patriots Program of 2014.
Snake first encountered Jonathan and the rest of the Rat Patrol squad during the events of Liquid Sun. After Snake was held up by Johnny, he was able to overpower the bumbling soldier and snatch his weapon, to which Jonathan, Ed, and Meryl Silverburgh, the team's commander, sprung to action. Shortly after, Rat Patrol and Snake were accosted by a battalion of Liquid Ocelot's personal army, the FROGS. During the battle, Jonathan made impressive use of his enormous size and power, along with his grenade launcher fitted to his XM8, while Snake and the others assisted in the escape. Making their way out of the mansion rendezvous point, they eventually convened in an underground parking garage before going their separate ways.
Later, during the events of Third Sun, Jonathan and the rest of the Rat Patrol attempted to arrest Liquid Ocelot, resulting in his deactivation of the SOP system, causing their weapons to seize up and not fire. Jonathan and Ed were both badly injured when the Rat Patrol's boat capsized, however they were able to survive.
After Liquid Ocelot was defeated and peace was restored, Jonathan acted as the best man in Meryl and Johnny's wedding.
